meaning of "flattened"

[Eway]

My dictionary says "flatten" could mean "to reduce to an even surface" or "to render stale". Which fits into the following contex?

"Stratum granulosum - 3-4 layers of cell thick consisting of flattened keratinocytes. At this level the cells are dying."

(Read from http://www.ridgesandfurrows.homestea...tion_skin.html)

[wsemajb]

Another annotation describes Squamous cells as being "identified histologically by the fact that they look flattened and thin..."

This would be describing their physical aspect - like a flat piece of paper.

Flattened in the sense of stale usually refers to taste, tone, or quality of something. Flat beer (lacking effervescence), flat singing, or even a flat mood. This is one of those words that never quite draws a clear distinction however, since flat beer lacks texture, flat singing might lack scale, and a flat mood is often indicated by a flat expression on ones face - each of which suggests something physically flat as well!