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For 8,000-to-10,000-pound creatures who spend all day on their feet and can live into their 40s, the consequence of that confinement was a painful middle age, marked by arthritis, cracked toenails and sore feet.
More: L.A. City Council should act to protect elephants - latimes.com
Does "can live into their 40s" refer to Life expectancy of elephants?
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