Cephalotus is peculiar in that

Maybo

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Seedlings grow slowly into low-growing plants that spread through creeping, branching underground rhizomes. Cephalotus is peculiar in that it produces two main forms of leaves. (The Savage Garden: Cultivating Carnivorous Plants by Peter D'Amato)

Does the bolded “that” refer to “Seedlings…rhizomes”?
 
@Maybo You could see it as a shortened form of the following sentence.

Cephalotus is peculiar in the sense that it produces two main types of leaves.
 
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